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A richly illustrated guide to lichens and their biology. Existing at the margins of life, lichens are the result of symbiotic relationships between fungi and photosynthesizing partners in the form of algae or cyanobacteria. Comprising more than twenty thousand species, lichens are pioneers in diverse ecosystems, colonizing virtually any surface and growing at almost any altitude. Found in rainforests, polar regions, deserts, and your backyard, lichens embody a paradox of toughness and sensitivity, surviving trips to space yet endangered by even the slightest environmental changes from industrial pollution here on Earth.